Describe the path that water takes through a plant from the roots to the leaves

The movement of plants from water to land has necessitated the development of internal mechanisms to supply all the parts of the plant with water. As discussed in Plant Classification, Vasular Tissues , tracheophytes (including virtually all terrestrial plants except for mosses and liverworts), have developed complex vascular systems that move nutrients and water throughout the plant body through "tubes" of conductive cells. The vascular tissues of these plants are called xylem and phloem. The xylem of vascular plants consists of dead cells placed end to end that form tunnels through which water and minerals move upward from the roots (where they are taken in) to the rest of the plant. Phloem, which is made up of living cells, carries the products of photosynthesis (organic nutrients) from the leaves to the other parts. The vascular system is continuous throughout the whole plant, even though the xylem and phloem are often arranged differently in the root than they are in the shoot.

The major mechanism by which water (along with dissolved materials) is carried upward through the xylem is called TATC (Transpiration-Adhesion-Tension-Cohesion). It should be noted that TATC, while supported by most scientists, is speculated but not proven to be at work in very tall trees. In this theory, transpiration, the evaporation of water from the leaf, is theorized to create a pressure differential that pulls fluids (held together by cohesion) up from the roots.

Water transport also occurs at the cellular level, as individual cells absorb and release water, and pass it along to neighboring cells. Water enters and leaves cells through osmosis, the passive diffusion of water across a membrane. In plants, water always moves from an area of higher water potential to an area of lower water potential. Water potential results from the differences in osmotic concentration (the concentration of solute in the water) as well as differences in water pressure (caused by the presence of rigid cell walls) between two regions. The relationship between the amount of dissolves solute and water potential is inverse: where there is a lot of dissolved solute the water potential is low.

Most of the water that a plant takes in enters through the root hairs. The water diffuses easily (and osmotically) into the root hairs because the concentration of dissolved materials in the plant's cellular cytoplasm is high. As discussed in Plant Classification, Root Hairs, there are two pathways through which water travels from the outside of the root to the core, where it is picked up by the xylem. The first of these pathways is the symplast, in which water moves across the root hair membrane and through the cells themselves, via channels that connect their contents. An alternate route for water is the apoplast, in which water travels along cell walls and through intercellular spaces to reach the core of the root. Once in the xylem, the water can be carried by TATC to all the other parts of the plant.

Overall, water is transported in the plant through the combined efforts of individual cells and the conductive tissues of the vascular system. Water from the soil enters the root hairs by moving along a water potential gradient and into the xylem through either the apoplast or symplast pathway. It is carried upward through the xylem by transpiration, and then passed into the leaves along another water potential gradient. In the leaf, some water is lost through evaporation from the stomata and the remaining fluid moves along a water potential gradient from the xylem into the phloem, where it is distributed along with the organic nutrients produced by photosynthesis throughout the plant.

Water and dissolved minerals from the soil enter plants through their roots. Many of the cells on the surface of the root contain root hairs. These projections increase the overall surface area increasing the ability of the root to take up more water.

The process by which water enters the root is osmosis.

Osmosis is the movement of water molecules from an area of high water concentration (weak/dilute solution) i.e. the soil to an area of low water concentration (strong/concentrated solutions) i.e. the plant cells through a partially permeable membrane (the cell membrane).

For an explanation of osmosis, click here!

Once water enters the root from the soil it travels to the xylem vessels in the middle of the root. The xylem vessels transport the water up through the stem and into the leaves of the plant. The leaves have a high concentration of dissolved minerals produced from photosynthesis and an overall low water concentration. Water is thus transferred from the xylem vessels to the cells in the leaves via osmosis.

The animation below shows water transport into the roots:


Agronomy and Horticulture Department

Title

Transpiration - Water Movement through Plants

Date of this Version

2005

Citation

Plant and Soil Sciences eLibrary (PASSeL) Lesson

Abstract

This lesson and its animation follow the journey of water through a plant from its uptake by roots to its evaporation from the leaf surface. How this journey is altered by plant characteristics such as stomata and cuticles as well as by changes in the environment will be described.

Overview

Transpiration is the loss of water from a plant in the form of water vapor. Water is absorbed by roots from the soil and transported as a liquid to the leaves via xylem. In the leaves, small pores allow water to escape as a vapor and CO2 to enter the leaf for photosynthesis. Of all the water absorbed by plants, less than 5% remains in the plant for growth and storage following growth. This lesson will explain why plants lose so much water, the path water takes through plants, how plants might control for too much water loss to avoid stress conditions, and how the environment plays a role in water loss from plants.

Objectives

At the completion of this lesson, students will be able to:

  1. Define transpiration and explain why it occurs in plants.
  2. Follow the pathway that water takes through plants from root uptake to evaporation at leaf cell surfaces.
  3. Describe how the driving force for water movement and any resistances to its flow through the plant are the two major components controlling rates of transpiration.
  4. Describe how environmental conditions alter rates of transpiration.
  5. Explain how the plant is able to alter rates of transpiration.

Modules:

  • Lesson home
  • Transpiration - Overview and Objectives
  • Transpiration - Introduction
  • Transpiration - What and Why?
  • Transpiration - What Controls Rates of Transpiration?
  • Transpiration - Major Plant Highlights
  • Transpiration - Factors Affecting Rates of Transpiration
  • Transpiration - Summary
  • Transpiration - References
  • Glossary
  • Videos

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What pathway does water take through a plant?

Water from the soil enters the root hairs by moving along a water potential gradient and into the xylem through either the apoplast or symplast pathway. It is carried upward through the xylem by transpiration, and then passed into the leaves along another water potential gradient.

What is the process of water passing through the plant from the root to the leaves and out through the stomata?

Transpiration is a process that involves loss of water vapour through the stomata of plants. The loss of water vapour from the plant cools the plant down when the weather is very hot, and water from the stem and roots moves upwards or is 'pulled' into the leaves.

How does water move through a plant step by step?

1-Water is passively transported into the roots and then into the xylem. 2-The forces of cohesion and adhesion cause the water molecules to form a column in the xylem. 3- Water moves from the xylem into the mesophyll cells, evaporates from their surfaces and leaves the plant by diffusion through the stomata.